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Holly and Poppy O'Hair are twins, but it's always been easy to tell these sisters apart. Because she is destined to be the next Rapunzel, Holly has always had the longest locks at Ever After High, while Poppy, whose fairytale destiny is up in the air, wears her hair short to express her individuality. But when Poppy's hair begins growing longer and longer, while Holly's hair gets shorter and shorter, it seems as if the sisters might have to reveal the secret they've been keeping about their destinies. Can they turn this bad hair day around, or will their fairytale ending be a fairy-fail?